(adj.) (sing. echina, pl. echinae) ( Wodehouse, 1928) Describing pollen and spores with an ornamentation comprising spines longer than 1µm. Comment: (Echinate should only be used in the adjectival form.) ( Erdtman (1952) recommended the term spinose, but in his usage spines were defined as longer than 3µm and smaller features as spinules (adj. spinulose). See also: microechinate, spine, spinule.